MODELING ELK & DEER POPULATION DYNAMICS IN IDAHO
IDAHO DEPARTMENT OF FISH AND GAME Steven M. Huffaker, Director Project W-160-R-32 Subproject 55-2 Progress Report
MODELING ELK & DEER POPULATION DYNAMICS IN IDAHO
July 1, 2004 to June 30, 2005
By: J. A. Manning Graduate Student University of Idaho E. O. Garton Professor of Wildlife Resources University of Idaho
Peter Zager Principal Wildlife Research Biologist
October 2005 Boise, Idaho
Findings in this report are preliminary in nature and not for publication without permission of the Director of the Idaho Department of Fish and Game.
The Idaho Department of Fish and Game adheres to all applicable state and federal laws and regulations related to discrimination on the basis of race, color, national origin, age, gender, or handicap. If you feel you have been discriminated against in any program, activity, or facility of the Idaho Department of Fish and Game, or if you desire further information, please write to: Idaho Department of Fish and Game, PO Box 25, Boise, ID 83707; or the Office of Human Resources, U.S. Fish and Wildlife Service, Department of the Interior, Washington, DC 20240.
This publication will be made available in alternative formats upon request. Please contact the Idaho Department of Fish and Game for assistance.
TABLE OF CONTENTS
MODELING ELK AND DEER POPULATION DYNAMICS IN IDAHO...................................1 ABSTRACT...............................................................................................................................1 INTRODUCTION .....................................................................................................................1 STUDY AREA ..........................................................................................................................2 METHODS ................................................................................................................................2 Equilibrium Densities ..........................................................................................................2 Annual Variability in Snow Depth and Summer Forage .....................................................3 Effects of Density Dependence, Inter-specific Competition, Winter Snow, Summer Forage, and Harvest on Population Growth.........................................................................4 RESULTS AND DISCUSSION ................................................................................................5 Equilibrium Densities ..........................................................................................................5 Annual Variability in Snow Depth and Summer Forage .....................................................6 Effects of Density Dependence, Inter-specific Competition, Winter Snow, Summer Forage, and Harvest on Population Growth.........................................................................7 MANAGEMENT IMPLICATIONS .........................................................................................7 LITERATURE CITED ..............................................................................................................8
LIST OF TABLES
Table 1. Variables used to construct linear regression models for predicting snowfall from daily DAYMET precipitation and temperature data......................................................................17 Table 2. Variables used to construct linear regression models for predicting snow depth from daily DAYMET precipitation and temperature data.............................................................17 Table 3. Variables used to construct linear regression models for predicting their singular and additive effects on population growth rates (r). ......................................................................18 Table 4. Linear regression models developed to predict monthly snowfall from daily DAYMET precipitation and temperature data in Idaho. ...............................................................18 Table 5. Linear regression models developed to predict monthly snow depth from snowfall in Idaho. .........................................................................................................................................19 Table 6. Linear regression models developed to predict singular and additive effects of density dependence, inter-specific competition, winter snow, summer forage, and harvest on population growth on population growth rates (r)....................................................................19
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TABLE OF CONTENTS (Continued)
LIST OF FIGURES
Figure 1. Study areas and ecoregions. ...........................................................................................11 Figure 2. Equilibrium densities of mule deer in GMUs (A) 11, (B) 21, and (C) 36B...................12 Figure 3. Equilibrium density of elk GMU 36B. ...........................................................................13 Figure 4. Predicted and observed snowfall in the Region 1, Elk River Ranger Station Snotel site, 1980-2003. ..................................................................................................................13 Figure 5. Predicted snowfall and snow depth in the Region 1, Elk River Ranger Station Snotel site, 1980-2003. ..................................................................................................................14 Figure 6. Mean NDVI in mule deer summer ranges in GMUs 11, 21, and 36B. ..........................14 Figure 7. Standard deviation of NDVI in mule deer summer ranges in GMUs 11, 21, and 36B. ................................................................................................................................................15 Figure 8. NDVI-values in mule deer summer ranges in GMUs (A) 11, (B) 21, (C) 36B, 1989................................................................................................................................................16
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PROGRESS REPORT STATEWIDE WILDLIFE RESEARCH
STATE:
Idaho
PROJECT TITLE: Modeling Large-Scale Elk
PROJECT:
W-160-R-32
and Deer Population
PROJECT NO.: 55
Dynamics in Idaho
SUBPROJECT: 2
PERIOD COVERED: July 1, 2004 to June 30, 2005
MODELING ELK AND DEER POPULATION DYNAMICS IN IDAHO
Abstract
Rocky Mountain elk and deer populations continue to exhibit large-scale changes in Idaho and throughout the western states. The preliminary results presented here are part of a larger study initiated to study the effects of competition and other factors on the dynamics of elk, mule deer, and white-tailed deer populations in order to predict population responses to various inter- and intra-specific factors. Here, we present estimates of mule deer and elk equilibrium densities and results from a model that predicts snow depth intended for use to estimate inter-annual changes in winter severity and amounts of winter range for mule deer and elk. We also demonstrate an application of satellite imagery to index forage quality in mule deer and elk summer ranges. Lastly, we show relative effects of competition and habitat condition on mule deer and elk in selected areas of Idaho.
Introduction
Rocky Mountain elk (Cervus elaphus nelsoni), mule deer (Odocoileus hemionus), and whitetailed deer (Odocoileus virginianus) populations extensively overlap throughout western North America. In Idaho and throughout the western states, their populations are experiencing largescale changes (Unsworth et al. 1995), and numerous intrinsic and extrinsic factors may influence such population fluctuations. Combined, these factors may have confounding effects on the fluctuations of deer and elk populations, which complicate our ability to predict the effects of management decisions. Combining these spatially and temporally variable factors in a predictive model that accounts for relative and interactive effects, including intra- and inter-specific competition, may provide accurate predictions of management decisions like harvest limits or predator control. Such models will assist wildlife managers in maintaining productive deer and elk herds at the regional level.
For analyses of population dynamics that include competition, it is useful to describe the range of population sizes of 2 species that results in one maintaining a zero population growth (r = 0) or equilibrium density (K) (Williams et al. 2002). "Ecological" carrying capacity has been defined as the size of a population when it is at equilibrium with its food supply (Caughley 1979), and can be derived from empirical relationships between population growth rate and population size (Houston 1982). Boyce (1990) and Boyce and Merrill (1991) relaxed the equilibrium assumption for elk by effectively making K a function of variable weather. Merrill and Boyce (1991) took
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